Moles were Talking Beasts that were smart, intelligent and peaceful animals. They were also excellent diggers and gardeners.
Moles were summoned by Aslan's call after Narnia's creation, and were later seen at the coronation of Frank I and his wife Helen, Narnia's first King and Queen.
Moles were also troops in Aslan's Army where they fought the White Witch's army in the First Battle of Beruna, in The Lion, the Witch, and the Wardrobe. One Mole, named Lilygloves, who was the chief, planted an orchard tree of fresh apples for the Pevensies.
In Prince Caspian, many Moles served and fought with the Old Narnians, with Caspian X and the Pevensies, against King Miraz and the Telmarine army in the Second Battle of Beruna. One Mole named Clodsley Shovel and his Moles dug up turf for the Dryads to eat.
Moles were among the talking beasts conversing with Shift the ape about Aslan and his whereabouts in The Last Battle. A pair of moles along with some mice and a rabbit gave food to and cleaned Tirian after he was tied to a tree by the Calormenes, with the moles carrying the bags holding the food and supplies. One of them swore to do it again despite the punishment if discovered, while the other one hushed and told him to be careful. More moles came to assist the king in the battle at Stable Hill and also assisted the mice and squirrels in freeing the talking horses.